by Samuel Foley Lipid membranes are fundamentally important structures in all branches of life. Not only do they form the boundary of living cells, they also perform a multitude of roles within them. To deeply understand these fluid membranes and their properties is then essential for explaining the mechanics of life at these scales. An important property of biological membranes which has recently moved to the forefront of research is the asymmetry of their constituent layers, as discussed in a.