Upwelling intensity structures free-living and particle-associated bacterial communities in an eastern boundary upwelling system

Eastern Boundary Upwelling Systems (EBUS) are among the most productive oceanic regions, driven by the upwelling of cold, nutrient-rich, and low-oxygen waters. These systems are increasingly affected by climate change, with intensified upwelling activity in frequency, strength, and duration, potentially reshaping microbial communities and their ecosystem functions. Furthermore, upwelling activity is not only subject to long-term climate change but also responds to natural variability across mult