New genera of black fungus gnats (Diptera: Sciaridae) from China―Part 2: Genus Cardiosciara Shah, Heller & Huang gen. nov.

Cardiosciara Shah, Heller & Huang gen. nov., formerly included within Cratyna Winnertz, 1867, is described and distinguished based on clear morphological and genetic differences. Diagnostic features of the new genus include long maxillary palpi, a setose posterior pronotum, an undifferentiated tibial organ, bare posterior wing veins, and male genitalia with a distinctive tegmen that is deeply notched apically with a dark, often heart-shaped overlapping structure. Four new species are describ